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Gaga Monstraparva
Scientists ________ Duke University ________ the US named a new genus of ferns discovered throughout the Americas ________ pop superstar Lady Gaga. When examining the ferns' DNA, the scientists spotted the base pair sequence G-A-G-A - just like the singer. They noted the ferns display gender fluidity, a major theme ________ Lady Gaga's work.
(Adapted from http://blog.oxforddictionaries.com)
Which is the correct way to complete the paragraph below?